shade-intolerantCOCA²⁸⁰¹³⁶⁺⁵BNC²³⁶²¹³⁺¹ 例句 In general, our experiments suggested that non-dormant seed and shade-intolerant seedlings of L. formosana were key factors limiting seedling regeneration under canopies of Formosan sweetgum forest. 因此枫香林下弱的光环境和种子的低温萌发特性是影响枫香幼苗更新的重要因子。 plant-ecology